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Sands, A.F.; Glöer, P.; Gürlek, M.E.; Albrecht, C.; Neubauer, T.A. (2020). A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution. 96(1): 25-66.
366257
10.3897/zse.96.48312 [view]
Sands, A.F.; Glöer, P.; Gürlek, M.E.; Albrecht, C.; Neubauer, T.A.
2020
A revision of the extant species of <em>Theodoxus</em> (Gastropoda, Neritidae) in Asia, with the description of three new species
Zoosystematics and Evolution
96(1): 25-66
Publication
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2020-02-03 13:22:57Z
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2022-08-23 06:42:30Z
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Nerita philippii Récluz, 1841 accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Neritina (Neritaea) gombaulti Pallary, 1939 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina (Neritaea) homsensis Pallary, 1939 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina (Neritaea) homsensis var. major Pallary, 1939 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina (Neritaea) homsensis var. minor "Pallary, 1939" accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina (Neritaea) jordani var. aberrans Dautzenberg, 1894 accepted as Neritina jordani G. B. Sowerby I, 1836 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina (Neritaea) jordani var. turris Mousson, 1861 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina (Neritaea) mesopotamica Mousson, 1874 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina (Theodoxia) jordani var. aberrans Dautzenberg, 1894 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina africana Reeve, 1856 accepted as Theodoxus africanus (Reeve, 1856) (additional source)
Neritina anatensis G. B. Sowerby II, 1849 accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Neritina anatolica var. mesopotamica E. von Martens, 1874 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina belladonna Mousson, 1854 accepted as Theodoxus anatolicus (Récluz, 1841) (source of synonymy)
Neritina bellardii Mousson, 1854 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina brauneri Lindholm, 1908 accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Neritina callosa Deshayes, 1835 accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Neritina cinctella E. von Martens, 1874 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina elongatula Morelet, 1845 accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Neritina euphratica Mousson, 1874 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina euxina Clessin, 1885 accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Neritina euxinus Clessin, 1885 accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Neritina fluviatilis var. abrauensis Milaschewitsch, 1913 accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Neritina guadianensis Morelet, 1845 accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Neritina heldreichi E. von Martens, 1879 accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Neritina heldreichi var. graeca Westerlund, 1886 accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Neritina hidalgoi Crosse, 1880 accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Neritina inquinata Morelet, 1845 accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Neritina jordani var. turris Mousson, 1861 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina karasuna Mousson, 1874 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina liturata Eichwald, 1838 accepted as Theodoxus major Issel, 1865 (source of synonymy)
Neritina macrii var. michoni [sic] accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina meridionalis var. mesopotamica Mousson, 1874 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina mesopotamica E. von Martens, 1874 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina mesopotamica Mousson, 1874 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina michonii Bourguignat, 1852 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina nilotica Reeve, 1855 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina orontis Blanckenhorn, 1897 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina ponsoti Pallary, 1930 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Neritina saulcyi Bourguignat, 1852 accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Neritina schultzii Grimm, 1877 accepted as Theodoxus major Issel, 1865 (source of synonymy)
Neritina velascoi Graells, 1846 accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Neritina violacea Morelet, 1845 accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxia ghigii Gambetta, 1929 accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Theodoxus (Neritaea) euphraticus (Mousson, 1874) accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Theodoxus (Neritaea) jordani tricarinatus Schütt, 1993 † accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Theodoxus (Neritaea) jordani var. bicarinatus Picard, 1934 † accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Theodoxus (Neritaea) jordani var. unicarinatus Picard, 1934 † accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Theodoxus (Neritaea) octagonus Eichhorst, 2016 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Theodoxus (Neritaea) pliocostulatus Schütt, 1993 † accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Theodoxus (Neritaea) varius (Rossmässler, 1835) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus (Neritaea) varius callosus (Deshayes, 1835) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus (Neritaea) varius varius (Rossmässler, 1835) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus (Ninnia) schultzi [sic] accepted as Theodoxus major Issel, 1865 (source of synonymy)
Theodoxus (Ninnia) schultzi var. jukovi V. P. Kolesnikov, 1947 accepted as Theodoxus major Issel, 1865 (source of synonymy)
Theodoxus (Theodoxus) brauneri Lindholm, 1908 accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Theodoxus (Theodoxus) callosus (Deshayes, 1835) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus (Theodoxus) elongatulus (Morelet, 1845) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus (Theodoxus) euxinus (Clessin, 1885) accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Theodoxus (Theodoxus) guadianensis (Morelet, 1845) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus (Theodoxus) heldreichi (E. von Martens, 1879) accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Theodoxus (Theodoxus) hidalgoi (Crosse, 1880) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus (Theodoxus) pallasi Lindholm, 1924 accepted as Theodoxus major Issel, 1865 (source of synonymy)
Theodoxus (Theodoxus) pallasi var. nalivkini V. P. Kolesnikov, 1947 accepted as Theodoxus major Issel, 1865 (source of synonymy)
Theodoxus (Theodoxus) saulcyi (Bourguignat, 1852) accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Theodoxus (Theodoxus) velascoi (Graells, 1846) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus africanus (Reeve, 1856) (additional source)
Theodoxus altenai Schütt, 1965 (basis of record)
Theodoxus anatolicus (Récluz, 1841) (basis of record)
Theodoxus astrachanicus Starobogatov, 1994 accepted as Theodoxus major Issel, 1865 (source of synonymy)
Theodoxus brauneri (Lindholm, 1908) accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Theodoxus callosus (Deshayes, 1835) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus cinctellus (E. von Martens, 1874) accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Theodoxus doriae Issel, 1865 accepted as Theodoxus pallidus (Dunker, 1861) (source of synonymy)
Theodoxus doriae var. obscura H. E. J. Biggs, 1937 accepted as Theodoxus pallidus (Dunker, 1861) (source of synonymy)
Theodoxus elongatulus (Morelet, 1845) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus euphraticus (Mousson, 1874) accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Theodoxus euxinus (Clessin, 1886) accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Theodoxus guadianensis (Morelet, 1845) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus gurur Sands & Glöer, 2020 (original description)
Theodoxus heldreichi (E. von Martens, 1879) accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Theodoxus heldreichi fluvicola Schütt & Şeşen, 1992 accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Theodoxus heldreichi heldreichi (E. von Martens, 1879) accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Theodoxus hidalgoi (Crosse, 1880) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus lituratus (Eichwald, 1838) accepted as Theodoxus major Issel, 1865 (source of synonymy)
Theodoxus macri (G. B. Sowerby II, 1849) (basis of record)
Theodoxus major Issel, 1865 (basis of record)
Theodoxus niloticus (Reeve, 1855) accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Theodoxus octagonus Eichhorst, 2016 accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Theodoxus orontis (Blanckenhorn, 1897) accepted as Theodoxus jordani (G. B. Sowerby I, 1836) (source of synonymy)
Theodoxus pallasi Lindholm, 1924 accepted as Theodoxus major Issel, 1865 (source of synonymy)
Theodoxus pallidus (Dunker, 1861) (basis of record)
Theodoxus sarmaticus (Lindholm, 1901) (additional source)
Theodoxus saulcyi (Bourguignat, 1852) accepted as Theodoxus fluviatilis (Linnaeus, 1758) (source of synonymy)
Theodoxus schultzii (Grimm, 1877) accepted as Theodoxus major Issel, 1865 (source of synonymy)
Theodoxus syriacus (Bourguignat, 1852) (basis of record)
Theodoxus varius (Rossmässler, 1835) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus varius callosus (Deshayes, 1835) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus varius varius (Rossmässler, 1835) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus velascoi (Graells, 1846) accepted as Theodoxus baeticus (Lamarck, 1822) accepted as Theodoxus meridionalis (R. A. Philippi, 1836) (source of synonymy)
Theodoxus velox V. Anistratenko, 1999 (basis of record)
Holotype NBCN RMNH.MOL.342197, identified as Theodoxus gurur Sands & Glöer, 2020
Holotype NBCN RMNH.MOL.342200, identified as Theodoxus wesselinghi Sands & Glöer, 2020
Holotype NBCN RMNH.MOL.342208, identified as Theodoxus wilkei Sands & Glöer, 2020
 Synonymy

Probably synonymous with either Theodoxus fluviatilis or T. velox[details]

 Type designation

Sakarya River, Çayköy, Bilecik, Turkey [details]

 Type locality

Suyunbaşı spring complex, Ayrancılar, İzmir, Turkey [details]

 Type locality

Çifteler spring, Çifteler, Eskişehir, Turkey [details]

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