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Betekhtina, O. A. (1975). Faunisticheskaya kharakteristika Izykhskogo razreza Minusinskogo basseyna [Faunal characteristics of the Izykh section of the Minusinsk basin]. In: Paleontologiya, stratigrafiya i paleobiogeografiya devona i karbona Sibiri [Paleontology, stratigraphy and paleobiogeography of the Devonian and Carboniferous of Siberia]. Akademiya Nauk SSSR, Sibirskoye Otdeleniye, Trudy Instituta Geologii i Geofiziki [Academy of Sciences of USSR, Siberian Branch, Transactions of the Institute of Geology and Geophysics]. 259: 28-35, pl. 5-6.
Betekhtina, O. A. (1975). Faunisticheskaya kharakteristika Izykhskogo razreza Minusinskogo basseyna [Faunal characteristics of the Izykh section of the Minusinsk basin]. In: Paleontologiya, stratigrafiya i paleobiogeografiya devona i karbona Sibiri [Paleontology, stratigraphy and paleobiogeography of the Devonian and Carboniferous of Siberia]. Akademiya Nauk SSSR, Sibirskoye Otdeleniye, Trudy Instituta Geologii i Geofiziki [Academy of Sciences of USSR, Siberian Branch, Transactions of the Institute of Geology and Geophysics]. 259: 28-35, pl. 5-6.